top biology phd programs

• The Best Universities for PhDs in Biological Sciences in the USA (2024)

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Why Study Biological Sciences in the USA

There are many reasons why someone might want to study Biological Sciences at a university in the USA . Here are just a few:

• From Marine Biology to Molecular Biology, the USA offers a broad spectrum of research topics in Biological Sciences. Candidates can align themselves with any niche that captivates their interest or fuels their academic curiosity.
• US institutions are home to highly esteemed faculty who have made significant contributions to the field of Biological Sciences. Their extensive knowledge and experience offer students an enriched learning experience.
• The strong ties between academic institutions and biotechnology or pharmaceutical industries in the USA open opportunities for internships, collaboration, and applied research prospects.
• Universities in the USA attract international students from all corners of the globe creating diverse educational settings that enrich both cultural understanding as well as academics. This multicultural environment exposes students to different perspectives within their field of study.

The Best Universities for Biological Sciences in the USA

The following tables give the 10 top universities in the USA for Biological Sciences , according to global and local university rankings. It can show you which American universities are amongst the best in the world - and help you compare institutions on an international level.

This information is based on the latest rankings tables, researched and published by Times Higher Education , QS and Academic Ranking of World Universities (ARWU) .

Each ranking system uses its own methodology, with different factors having more or less influence on a university's result.

Our guide has more information on how to use international rankings to decide on the best research universities for PhD study .

What should I know about the Times Higher Education rankings?

The Times Higher Education rankings are strong in academic focus and diverse teaching metrics, but do not include employer-specific metrics. Additionally, the rankings may not include all specialist institutions.

What should I know about the QS rankings?

The QS World University Rankings are designed to meet the needs of prospective students, with more weight given to student-centric metrics such as staff/student ratio, international recruitment and employer opinion. The rankings are balanced between qualitative and quantitative data, but give less weight to research than some other rankings.

What should I know about the ARWU rankings?

The ARWU rankings reflect the presence of elite academics and the future academic success of graduates. However, they do not directly assess the quality of education at a university or take into account other aspects of university performance.

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Graduate Admissions

Helpful guides for applying to the biology phd program, tips for applying to the cmob track, tips for applying to the eco/evo track.

The PhD program trains students for careers in research and teaching. In addition to a wide range of courses, the PhD program offers informal but intensive instruction in laboratories and seminars, characterized by close contact between students and their faculty advisors. Biology is one of fourteen home programs  in the  Biosciences,  which span the School of Medicine and the School of Humanities and Sciences.

How to Apply

For the 2023-2024 PhD application season, application processes will be run through existing Biology (School of Humanities & Sciences)  and the Oceans Department(Stanford Doerr School of Sustainability). 

Previously part of the Biology Department , Hopkins Marine Station is now part of the Oceans Department within the Stanford Doerr School of Sustainability (SDSS).  Several faculty took appointments in Oceans, while others chose to stay in Biology; still others have joint appointments.  Additionally, several faculty members in Ecology/Evolutionary Biology now have joint appointments within Biology and SDSS. Based on these changes, the following information will provide further guidance for the application process. 

• Faculty with 100% appointments in Biology, will only recruit through Biology.  The Biology application will be used.
• Ecology/Evolutionary Biology faculty with 50/50 appointments in both Biology and SDSS will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Ecology/Evolutionary faculty with joint Biology/SDSS appointments, you should reach out to them directly regarding the appropriate application track to choose. 
• 100% Oceans faculty will only be recruiting students through the Earth Systems Science application.
• Oceans faculty appointed 50/50 in Biology and SDSS (Oceans Department), will recruit students through either Biology or Earth Systems Science .  If you are interested in working with Oceans faculty with joint Biology/SDSS appointments, you should reach out to the individual faculty members you hope to work with regarding the appropriate application and track to choose. 

Faculty Affiliated with the Stanford Doerr School of Sustainability

Oceans Department:

Fio Micheli 100% 

Jeremy Goldbogen 100%

Giulio De Leo 100%

Larry Crowder 100%

Steve Palumbi 50% Oceans/50%Biology

Barbara Block 50% Oceans/50% Biology

*Note Chris Lowe will remain 100% in Biology

Ecology/Evolution

Liz Hadly 50% SDSS/50%Biology

Rodolfo Dirzo 50% SDSS/50%Biology

Tad Fukami 50% SDSS/50%Biology

Kabir Peay 50% SDSS/50%Biology

Chris Field 50% SDSS/50% Biology

Gretchen Daily 50% SDSS/50% Biology

Application and Deadline

Stanford Biology PhD Program applications are made through Graduate Admissions . The application deadline for Autumn Quarter 2024 matriculation is December 5, 2023 at 11:59pm pst.  The application for the Autumn 2024 cohort will be available in September 2023.  Please review the Graduate Admissions website prior to starting your application. Applications are considered for admission in Autumn Quarter only. 

Application Status

To check your application status,  click here to Visit Your Status Page .  

Requirements

• U.S. Bachelor's degree or its foreign equivalent
• Stanford Graduate application
• Non-refundable application fee of $125 (or fee waiver found here ) 
• Statement of Purpose
• Three Letters of Recommendation (a maximum of four letters will be accepted)
• Transcripts - Unofficial transcripts from all universities and colleges you've attended for one year or more must be uploaded to the online application.  Graduate Admissions only requires admitted applicants who accept the offer of admission to submit official transcripts that shows their degree conferral. Please do not send or have sent any official transcripts to us at this time.
• TOEFL Test of English as a Foreign Language (required for non-native English speakers)
• A current Curriculum Vitae/Resume

Knight-Hennessy Scholars

Join dozens of  Stanford School of Humanities and Sciences students  who gain valuable leadership skills in a multidisciplinary, multicultural community as  Knight-Hennessy Scholars  (KHS). KHS admits up to 100 select applicants each year from across Stanford’s seven graduate schools, and delivers engaging experiences that prepare them to be visionary, courageous, and collaborative leaders ready to address complex global challenges. As a scholar, you join a distinguished cohort, participate in up to three years of leadership programming, and receive full funding for up to three years of your PhD studies at Stanford. Candidates of any country may apply. KHS applicants must have earned their first undergraduate degree within the last seven years, and must apply to both a Stanford graduate program and to KHS. Stanford PhD students may also apply to KHS during their first year of PhD enrollment. If you aspire to be a leader in your field, we invite you to apply. The KHS application deadline is October 11, 2023. Learn more about  KHS admission .

Biology PhD Program General Information

Diversity, inclusion, equity, and belonging.

The Department of Biology recognizes that the Supreme Court issued a ruling in June 2023 about the consideration of certain types of demographic information as part of an admission review. All applications submitted during upcoming application cycles will be reviewed in conformance with that decision. The Department of Biology welcomes graduate applications from individuals with a broad range of life experiences, perspectives, and backgrounds who would contribute to our community of scholars. Review of applications is holistic and individualized, considering each applicant’s academic record and accomplishments, letters of recommendation, and admissions essays in order to understand how an applicant’s life experiences have shaped their past and potential contributions to their field.Please see the websites below for more resources.

• Diversity and Engagement in Biosciences PhD Programs
• Vice Provost for Graduate Education
• Biology Equity and Belonging
• Research Sites
• Faculty Profiles and Labs
• Inquiries about the PhD in Biology should be directed to  biologyadmissions [at] lists.stanford.edu (biologyadmissions[at]lists[dot]stanford[dot]edu)

2024 Best Biological & Biomedical Sciences Doctor's Degree Schools

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Biological & Biomedical Sciences by Region

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Biological and Biomedical Sciences

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Biological and Biomedical Sciences (BBS)  is an area of study within the Division of Medical Sciences, an administrative unit based at Harvard Medical School that coordinates biomedical PhD activities at the Longwood Medical Area. Students who study in BBS receive a PhD in medical sciences. Prospective students apply through the Harvard Kenneth C. Griffin Graduate School of Arts and Sciences (Harvard Griffin GSAS). In the online application, select  “Division of Medical Sciences” as your program choice and select "Biological and Biomedical Sciences" in the area of study menu.

This program trains you in the biosciences starting with core training in contemporary genetics, biochemistry, and molecular, cellular, and mechanistic biology. You can customize your curriculum to align with your research interests.

You will have access to a vast wealth of resources, including more than 700 faculty, top scientists, core facilities and hospitals, cutting-edge research, and an extensive alumni network. You will be in close proximity to top nationwide hospitals and research centers and more than 130 biotech, biopharma, and pharmaceutical companies that have biological and biomedical sciences (BBS) alumni on staff.

Examples of specific student projects include “Development of nucleic acid detection methods for object provenance and viral diagnostics,” “Interrogating genetic diversity in Mycobacterium abscessus with transposon-sequencing,” and “Sexual dimorphism in hair follicle stem cells.”

Graduates of the program have secured faculty positions at such prestigious institutions as New York University, University of Minnesota, Harvard University, and University of Illinois. Others have gone on to careers with leading companies such as Novartis and the Broad Institute. Derrick Rossi, cofounder of Moderna, is a graduate of the BBS program.

Personal Statement

Standardized tests.

GRE General: Not Accepted GRE Subject: Not Accepted iBT TOEFL minimum score: 100 IELTS minimum score: 7

See list of Biological and Biomedical Sciences faculty

APPLICATION DEADLINE

Questions about the program.

Doctor of Philosophy (PhD) in Biology

The PhD in Biology is a research degree requiring graduate-level coursework, completion of a dissertation, and two semesters of participation in teaching (usually as a teaching fellow in laboratory or discussion sections of lecture courses led by Biology faculty). For most students, obtaining this degree typically involves five or more years of full-time study.

A summary of Biology PhD student expectations by year can be found here . Full details can be found in the Graduate Program Guide .

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program.

How to Apply    Frequently Asked Questions

Learning Outcomes

1. Demonstrate academic mastery in one of three areas of Biology: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.

2. Attain research expertise , including grant writing experience, and complete original research that advances a specific field of study within one of three broad subject areas represented in the department: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.

3. Attain teaching experience and expertise in one of three broad areas of Biology: Ecology, Behavior & Evolution; Neurobiology; or Cellular & Molecular Biology.

4. Attain the skills and qualifications needed for employment in an academic, government, or private sector position related to the life sciences.

Coursework Requirements

Students must complete 64 credits with a minimum grade point average of 3.0; at least 32 of these credits must be accrued from lecture, laboratory, or seminar courses. Students with prior graduate work may be able to transfer course credits.  See the  Graduate School of Arts & Sciences (GRS) Transfer of Credits policy  for more details. 

• 2 semesters of Progress in Research Seminars (2 credits each):

Cell & Molecular Biology:  BI 583 & BI 584 (CM section)

Ecology, Behavior & Evolution: BI 579 & BI 580

Neurobiology:   BI 583 & BI 584 (NEURO section)

• 1 semester grant-writing course (2 credits):

Cell & Molecular Biology:  BI 581

Ecology, Behavior & Evolution: BI 671

Neurobiology: BI 581

• 1 semester pedagogy course (1 credit): All Biology PhD students take  BI 697
• 1 quantitative course (3–4 credits): from a list of recommended courses .
• Additional courses vary by specific track below:

Cell & Molecular Biology

1. GRS BI 791/GRS BI 792 Graduate Rotation Credits (2 credits each/4 credits total)

2. GRS BI 753 Advanced Molecular Biology  (4 credits)

3. GRS MB 721 Graduate Biochemistry  (4 credits)

4. GRS BI 735 Advanced Cell Biology  (4 credits)

5. Two electives (8 credits, 500-level and above), see the Graduate Program Guide for recommendations

6. Research credits  (remaining credits)

Ecology, Behavior & Evolution

1. Six electives  (24 credits, 500-level and above), see the Graduate Program Guide for recommendations

2. Research credits  (remaining credits)

Ecology, Behavior & Evolution PhD candidate coursework is highly variable. Students, in consultation with advisors, develop a plan of coursework and research. Students are required to take a minimum of 32 credits of coursework. The remainder of the credits should be research.

Neurobiology

2. GRS BI 755 Cellular and Systems Neuroscience  (4 credits)

3. GRS BI 741 Neural Systems: Functional Circuit Analysis  (4 credits)

4. Four electives  (16 credits, 500-level and above), see the Graduate Program Guide for recommendations

5. Research credits  (remaining credits)

Teaching Requirement

The department requires a minimum of two semesters of teaching as part of the Doctor of Philosophy program. During the first semester of teaching, students are required to enroll in our first-year seminar course, GRS BI 697 A Bridge to Knowledge . The course provides guidance and training on pedagogy and other aspects of graduate school.

Qualifying Examination

The qualifying examination must be completed no later than six semesters after matriculation. In most graduate curricula in the department, this consists of a research proposal—often in the form of a grant application—which the student submits to their committee and subsequently defends in an oral presentation. In the  Cell & Molecular Biology and Ecology, Behavior & Evolution curricula , this is preceded by a comprehensive written examination testing the student’s general background from coursework.

Dissertation & Final Oral Examination

Candidates shall demonstrate their abilities for independent study in a dissertation representing original research or creative scholarship. A prospectus for the dissertation must be completed and approved by the readers, the Director of Graduate Studies, and the Biology Department Chair. Candidates must undergo a final oral examination in which they defend their dissertation as a valuable contribution to knowledge in their field and demonstrate a mastery of their field of specialization in relation to their dissertation. All portions of the dissertation and final oral examination must be completed as outlined in the  GRS General Requirements for the Doctor of Philosophy Degree . The results of the dissertation must be presented at a department colloquium.

Graduation Timeline

Forms and additional information about PhD graduation can be found on the GRS website .

9 – 12 months before proposed graduation date

• Dissertation Prospectus & approval form due to Graduate Program Specialist for departmental review and submission to GRS

Semester prior to your intended graduation cycle

• Intent to Graduate Form completed online

About 2 months before dissertation defense

• Meet with Graduate Program Specialist, Director of Graduate Studies, and Faculty Advisor to review requirements for defense
• Arrange for Special Service Appointments if committee members are not BU faculty
• Send first draft of dissertation to readers

Once defense date is confirmed with committee

• Reserve room(s) for public seminar and defense

At least three weeks prior to dissertation defense

• Schedule of Final Oral Exam with Abstract Approval due to GRS
• Properly formatted draft of dissertation submitted as PDF to [email protected]

At least two weeks prior to dissertation defense

• Send dissertation to all committee members

At least one week prior to dissertation defense

• Send program information to Graduate Program Specialist

See the Graduate Program Guide for final dates to submit dissertation to ETD

• Submit final dissertation to ETD (online submission)
• Complete electronic signature page and forward confirmation to Graduate Program Specialist along with a copy of your submitted dissertation

MS Degree (En Route to PhD)

Option one: A PhD student who has advanced to candidacy (as demonstrated by passing the PhD qualifying exam), and has completed 32 credits of graduate-level coursework (not including research) may apply to the Graduate School for an MS degree in Biology. This must be approved by the Director of Graduate Studies within the Biology Department. The student’s major professor will receive notification of this application process.

Option two: A PhD student who has taken, but has not advanced to candidacy based on the PhD qualifying examination, may still receive an MS degree. This student may receive a Coursework MS degree provided they have completed 32 credits of coursework (not including research credits). Alternatively, this student may receive a Scholarly Paper or Research Thesis MS degree if the written portion of the qualifying examination is adapted to ensure it is of sufficiently high quality for a MS degree, and approved by a majority of the qualifying exam committee and the Director of Graduate Studies.

Interdisciplinary Study Options

Biology PhD students have the option to participate in the Boston University Graduate Program in Urban Biogeoscience and Environmental Health (BU URBAN), the National Science Foundation Research Traineeship Program Understanding the Brain: Neurophotonics (NSF NRT UtB: Neurophotonics), and the Biogeoscience Advanced Graduate Certificate Program . These programs require separate applications in addition to the standard Biology PhD application; those interested in BU URBAN are encouraged pre-apply .

Time Limits

Officially, the PhD must be completed within seven years after the first registration for doctoral study. PhD degrees are conferred in either May, August, or January, as specified on the GRS website . In addition, the PhD candidacy expires after the fifth anniversary of passing the Qualifying Examination. Petitions to extend this deadline are possible at the discretion of the Director of Graduate Studies, the Department Chair, and the Dean of the Graduate School, and can be obtained from the Office of the Graduate School of Arts & Sciences.

The Biology Department guarantees support for five years for all PhD students, contingent on satisfactory performance in the program. PhD students are encouraged to apply for fellowships and grants at funding agencies. All domestic students should apply for NSF Graduate Research Fellowships in the Fall semester of their first or second year. 

Travel Grants may be available to assist students in their travel to professional scientific meetings; students presenting papers or posters on their research will receive first consideration.

Common Types of Funding:

Dean’s Fellowships: These are non-service fellowships allocated to first-year PhD students that do not have immediate teaching requirements.

Teaching Fellowships: These provide a stipend plus full tuition and fees for up to four full courses per semester plus a 2-credit teaching course. Teaching responsibilities usually require approximately 20 hours per week. Full or partial awards may be given.

Doctoral Research Fellowships: These awards are given to students who assist individual faculty with specific areas of research. These Research Fellowships provide a stipend and full tuition. The supervising faculty member determines the specific duties of the Research Fellow.

In addition to the above funding sources, several competitive Department awards and fellowships are available to graduate students in the Department of Biology.

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Stanford Cancer Institute

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Cancer Biology PhD Program

Established in 1978, the interdisciplinary Cancer Biology PhD Program is designed to provide graduate and medical students with the education and training they need to make significant contributions to the field of cancer biology. The program is led by Laura Attardi, PhD, and Julien Sage, PhD, and currently has over 60 participating faculty members representing a variety of specialties.

Coursework during the first year equips students with a broad understanding of the molecular, genetic, cellular biological and pathobiological aspects of cancer. By the beginning of the second year, students have chosen a research advisor and begun work on their dissertation projects. An annual program conference provides students with an opportunity to present their research findings, receive feedback and forge collaborations with faculty and fellow students. 

The Cancer Biology program also sponsors a postdoctoral scholar track in which accepted MD and PhD scholars pursue research under the guidance of a faculty mentor. 

Program Directors

Laura Attardi

Catharine and howard avery professor of the school of medicine and professor of genetics.

Julien Sage

Elaine and john chambers professor of pediatric cancer and professor of genetics.

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Thank you for taking the time to explore the University of Washington’s doctoral program in Biology. As you browse these pages, you will see that we value a diversity of people and approaches to solve the most important problems in Biology. Accordingly, the Department of Biology has an unusual breadth of top-notch faculty, postdocs, and students who study topics at spatial scales ranging from molecules to the entire planet.

Your individual scholarship and research are the highest priority while you are studying at the University of Washington. Through assistantships, assistance with grant proposals, and over a dozen departmental awards, our department works to open every avenue for you succeed in your research. And your opportunities aren’t limited to Biology. We encourage you to take advantage of the opportunity to collaborate with faculty throughout the University, as many of our faculty do.

These efforts are facilitated by Seattle’s position as a major intellectual hub. From cutting edge research foundations to the slopes of Mt. Rainier, the Pacific Northwest provides opportunities for biologists to enhance their learning at every turn. Additionally, the University of Washington has been the leading recipient of federal research funding—now more than $1 billion per year—among public universities for more than 20 years.

We look forward to receiving your application and discussing the exciting opportunities available to you.

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Ph.D. Program

The philosophy of the PhD program, along with the Affiliated Ph.D.  Program with the Salk Institute for Biological Studies, is to provide world-class research training in the basic biological sciences to equip a diverse group of trainees for a variety of scientific careers ranging from academia and industry to education, communication, or policy. Core principles of the program are to be student centered and attuned to the goals of the trainee.

The core curriculum focuses on development of core competencies and transferable skills in critical thinking, communication, and leadership. The first year prepares students for the core of the PhD program, the development of intellectual independence and creativity through original thesis research, guided by a thesis advisor and committee. Students have a high degree of flexibility in choice of thesis advisor through the rotation program. Throughout the program, there is strong emphasis on engaged mentoring through regular committee meetings, annual reports and Individual Development Plans.

As a central hub of the thriving San Diego biosciences community, the program maintains strong partnerships with other campus units and programs through joint faculty appointments, organized research units, and research collaborations, enabling a wide range of interdisciplinary opportunities . The mission is to conduct leading edge research in the basic biological sciences. Major areas of emphasis currently include structural biology, cell biology, developmental biology, neurobiology, immunology, microbiology, virology, plant biology, ecology, and evolutionary biology. Research in the School has emphasized studies using model organisms or in vitro mechanistic approaches, with human studies and clinical research concentrated in other departments or in the Health Sciences. Current and future areas of growth include quantitative biology, data science, and the biological consequences of climate change.

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• Commitment to quality mentorships, student mental health, and well-being
• Equity, Diversity and Inclusion as integral to program admissions and retention
• Open science practices valuing multiple research outputs and holistic assessment of scholarly excellence
• Communication and outreach as key aspects of scientific training

As a doctoral program embedded in a large undergraduate instructional unit, our approach incorporates substantial training in teaching methodology and best practices. Our philosophy remains that teaching and research are interdependent facets of engaged scholarship.

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So I'm sure this question had been asked many times before, but I didn't really find many satisfactory answers by searching so hopefully you all won't mind answering it one more time. I'm currently a year away from getting my BS in cellular/molecular biology and I'm pretty sure I want to go on to get my PhD. I recently read an article about Drew Endy 's research in synthetic biology at Stanford and decided that this is the kind of thing that I'd really love to do. I live in the bay area, so I actually went and walked around Stanford yesterday and met with a few members of the faculty to discuss their research and the biology program in general. I decided I'd probably like to apply to the bioengineering program (I really love math, physics, and computer science as well) over the biology program but of course even with my good grades and research experience I absolutely should not count on getting in there. I really haven't looked into too many other schools, but I've heard you should apply to around 10. I really love California and the bay area so I'd love to go someplace nearby. I've heard good things about Berkeley, UCSF, and UC Santa Cruz but I'm not aware of any good source that rates their biology programs specifically.

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John Chappell to direct Translational Biology, Medicine, and Health Graduate Program

Building on a decade of growth, the interdisciplinary research program prepares graduates for impactful careers in a dynamic biomedical landscape.

• Whitney Slightham

29 Jul 2024

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John Chappell , associate professor at the Fralin Biomedical Research Institute at VTC , has been appointed to direct Virginia Tech’s Translational Biology, Medicine, and Health (TBMH) Graduate Program, effective July 1. 

Chappell will take over for Steven Poelzing , professor at the Fralin Biomedical Research Institute, who has led the program for eight years. Michael Friedlander , Virginia Tech’s vice president for health sciences and technology and executive director of the Fralin Biomedical Research Institute, announced the appointment and praised Poelzing’s contributions.

“Dr. Poelzing has been instrumental in shaping the TBMH program into a nationally recognized leader in interdisciplinary biomedical research,” Friedlander said. “His legacy includes expanding the curriculum to foster a broader educational experience and significantly enhancing student success in securing prestigious fellowships and grants. We are deeply grateful for his dedication and visionary leadership.”

Poelzing, who is also a professor in the Department of Biomedical Engineering and Mechanics in the Virginia Tech College of Engineering, will continue to participate in the translational biology, medicine, and health program while turning his energy toward his research in the institute’s Center for Vascular and Heart Research.  

The program has seen steady growth under Poelzing’s leadership. In the past year alone, the program grew to 170 applicants from 112 in 2023 and boasts a doctoral degree completion rate that exceeds national averages. To date, translational biology, medicine, and health students have secured approximately $2 million in competitive national grant funding. Program alumni have pursued prestigious postdoctoral positions and fellowships at the National Institutes of Health, Mayo Clinic, Harvard University, Yale, MIT, and more, as well as leadership roles in industry and scientific policy institutions.

“I am thrilled to take on this new role and build on the solid foundation that Dr. Poelzing and the TBMH team have established,” said Chappell, who is also an associate professor in the College of Engineering’s Department of Biomedical Engineering and Mechanics . “My goal is to continue evolving the TBMH program to meet the changing landscape of biomedical research, ensuring our students are well-prepared for diverse career paths in academia, industry, and beyond. I look forward to working with our talented faculty and dedicated students to look ahead and drive innovation.”

Chappell brings extensive experience in student mentorship, recruitment, and program development. His commitment to student success and his deep understanding of the program’s interdisciplinary mission make him a natural fit to grow the program, according to Friedlander.

“Dr. Chappell’s passion for student training and his proven track record in leadership will be invaluable as we continue to develop and expand TBMH,” Friedlander said. “He listens to and advocates for students, and likewise they unanimously speak to his dedication and understanding. Overall, he has shown himself as a very effective ambassador for the program, and I am confident that under his guidance, TBMH will achieve new heights in biomedical research education.”

Chappell is known for fostering an inclusive and innovative learning environment. His vision includes expanding recruitment efforts to attract top-tier students and enhancing the program’s reputation for producing graduates who are not only skilled researchers but also leaders in the biomedical field.

He joined Virginia Tech in 2014 after completing a postdoctoral research fellowship at the University of North Carolina at Chapel Hill. He holds a doctoral degree and master’s degree in biomedical engineering and a bachelor’s degree electrical engineering from the University of Virginia.

Leigh Anne Kelley

540-526-2002

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The Unique Burial of a Child of Early Scythian Time at the Cemetery of Saryg-Bulun (Tuva)

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Pages:  379-406

In 1988, the Tuvan Archaeological Expedition (led by M. E. Kilunovskaya and V. A. Semenov) discovered a unique burial of the early Iron Age at Saryg-Bulun in Central Tuva. There are two burial mounds of the Aldy-Bel culture dated by 7th century BC. Within the barrows, which adjoined one another, forming a figure-of-eight, there were discovered 7 burials, from which a representative collection of artifacts was recovered. Burial 5 was the most unique, it was found in a coffin made of a larch trunk, with a tightly closed lid. Due to the preservative properties of larch and lack of air access, the coffin contained a well-preserved mummy of a child with an accompanying set of grave goods. The interred individual retained the skin on his face and had a leather headdress painted with red pigment and a coat, sewn from jerboa fur. The coat was belted with a leather belt with bronze ornaments and buckles. Besides that, a leather quiver with arrows with the shafts decorated with painted ornaments, fully preserved battle pick and a bow were buried in the coffin. Unexpectedly, the full-genomic analysis, showed that the individual was female. This fact opens a new aspect in the study of the social history of the Scythian society and perhaps brings us back to the myth of the Amazons, discussed by Herodotus. Of course, this discovery is unique in its preservation for the Scythian culture of Tuva and requires careful study and conservation.

Keywords: Tuva, Early Iron Age, early Scythian period, Aldy-Bel culture, barrow, burial in the coffin, mummy, full genome sequencing, aDNA

Information about authors: Marina Kilunovskaya (Saint Petersburg, Russian Federation). Candidate of Historical Sciences. Institute for the History of Material Culture of the Russian Academy of Sciences. Dvortsovaya Emb., 18, Saint Petersburg, 191186, Russian Federation E-mail: [email protected] Vladimir Semenov (Saint Petersburg, Russian Federation). Candidate of Historical Sciences. Institute for the History of Material Culture of the Russian Academy of Sciences. Dvortsovaya Emb., 18, Saint Petersburg, 191186, Russian Federation E-mail: [email protected] Varvara Busova  (Moscow, Russian Federation).  (Saint Petersburg, Russian Federation). Institute for the History of Material Culture of the Russian Academy of Sciences.  Dvortsovaya Emb., 18, Saint Petersburg, 191186, Russian Federation E-mail:  [email protected] Kharis Mustafin  (Moscow, Russian Federation). Candidate of Technical Sciences. Moscow Institute of Physics and Technology.  Institutsky Lane, 9, Dolgoprudny, 141701, Moscow Oblast, Russian Federation E-mail:  [email protected] Irina Alborova  (Moscow, Russian Federation). Candidate of Biological Sciences. Moscow Institute of Physics and Technology.  Institutsky Lane, 9, Dolgoprudny, 141701, Moscow Oblast, Russian Federation E-mail:  [email protected] Alina Matzvai  (Moscow, Russian Federation). Moscow Institute of Physics and Technology.  Institutsky Lane, 9, Dolgoprudny, 141701, Moscow Oblast, Russian Federation E-mail:  [email protected]

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When asked about doctoral students pursuing a policy career, all three groups we surveyed showed a preference for institutions with strong connections to Washington. Schools such as George Washington University, Georgetown University, and American University are ranked higher for students pursuing policy careers than for those hoping to enter academia. Conversely, institutions such as the University of Chicago and Massachusetts Institute of Technology are ranked lower for policy careers than academic ones. This shift highlights the importance of proximity to policy centers and the availability of practical engagement opportunities in shaping perceptions of a program’s value for a policy-oriented career in IR.

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Irene Entringer García Blanes is a senior project manager for the Teaching, Research, and International Policy Project at William & Mary. Twitter:  @EntringerIrene

Susan Peterson is the Wendy and Emery Reves professor of government and international relations at William & Mary.

Michael J. Tierney is the George and Mary Hylton professor of international relations and director of the Global Research Institute at William & Mary. Twitter:  @MikeTierneyIR

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